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The first isolation of guanine was reported in 1844 by the German chemist Julius Bodo Unger [de] (1819–1885), who obtained it as a mineral formed from the excreta of sea birds, which is known as guano and which was used as a source of fertilizer; guanine was named in 1846.
Cytosine was discovered and named by Albrecht Kossel and Albert Neumann in 1894 when it was hydrolyzed from calf thymus tissues.[4][5] A structure was proposed in 1903, and was synthesized (and thus confirmed) in the laboratory in the same year.
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